the expression refers to water's ability and predictability in always flowing to where it will be level. (well, in its liquid state anyway.)
water will not stay on the top of a hill if it can run downhill.
neither will it naturally defy gravity and move uphill on its own. neither does it bunch up in lumps -- the same quantity of water under the same conditions will flow to a calm state in the same place every time.
the expression means that 'things will naturally run their course', or 'people will naturally find their way'. depending on context, it can also mean that 'people will naturally do what is in their nature to do', no more, no less.
for example, say you think a person has gotten a promotion and can't do the job. i might say to you, "don't worry, water finds its own level", which means that i think that if this person isn't up to the job, it will become evident soon enough, and there is no reason to worry about it. it may also be true that he will be able to the job, and we should wait and see.
Water runs downhill, till it gets to the bottom.
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